MD, University of Iowa, Roy J. and Lucille A. Carver College of Medicine, Iowa City, Iowa
PhD, Molecular and Cellular Biology, University of Iowa, Iowa City, Iowa
BS, Biochemistry and Molecular Biology, Nebraska Wesleyan University, Lincoln, Neb.
Clinical Pathology, University of Iowa Hospitals & Clinics, Iowa City, Iowa
Molecular Genetic Pathology, University of Iowa Hospitals & Clinics, Iowa City, Iowa